What is the natural way that lion populations are controlled?
As with most predators, lion numbers would naturally (ie prior to losing enormous areas from their range) vary with the amount of available prey. A bumper year for wildebeeste, for instance, would mean a bumper year for lions, and therefore plenty of cubs surviving to adulthood. A failure of the rains, leading to reduced herbivore numbers, leads to fewer cubs surviving to maturity, and overall population numbers dropping off rapidly.
Hyenas eating the young,
No
or in times of drought did the males murder their offspring?
No. Lions kill (but don't eat) the offspring of a pride which they take over (one of the reasons that lion hunting has to be carefully controlled and thought through).
And can we encourage the natural method? ot have they been controlled for millenia by indigenous paople with spears and the need to have a rite of passage?
That misses the point entirely. Lion numbers across Africa are reducing catastrophically. We shouldn't be doing much to control their population, but should be doing what we can to increase supply of their prey, reduce the high level of poaching, and increase the amount of land available for them. Apex predators are a useful indicator of the health of the biomass "below" them in the food chain, and this park (which isn't a national park, but a private reserve formed from former farmland) clearly has some management issues.
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