Gosh... why does that whole article read like something written by someone with poor science literacy?
Even Darwin struggled to explain why we would evolve a response that lets others know that we have cheated or lied
Darwin didn't know a lot of things. Darwin did not know how the eye could evolve. Today we do. Darwin did not know the structure of genes. Now we do.
Modern science moved way beyond him, long ago. Why mention his name in such a way?
It's like saying: "Even Newton struggled to work out gravitational effects of three or more bodies at a time", as if that justifies calling it a mystery, today.
But, then the article answers its own question:
Noting that women blush more than men, neuroscientist V. S. Ramachandran of the University of California, San Diego, suggests that blushing might have evolved as a way for women to demonstrate their honesty to men and so enlist their help in rearing offspring.
I don't know if that is true, or not. But, now that we have a testable idea, why keep it on the list?
The emergence of Altruism has been well understood for a long time, as a way to maximize payoff in a group of seemingly selfish actors. The author clearly has not read anything on the subject since the 1950's or so.
The urge to kiss is not brought about by genes, so why do we find it so pleasurable to share saliva?
I bet genes does have something to do with it. Any isolated culture of humans will eventually converge on the act of kissing to show affection, demonstrating that genes will make a species more prone or less prone to do so. The author seems to be speaking from the point of view of someone who has done jack-squat research on the subject.
There
can be legitimate discussion on things we do not, yet, understand about humans. But, it is difficult to do so, when one is writing like a middle school student with no school library to borrow books from.