The Chimpanzee & Human Communication Institute (CHCI) at Central Washington University in Ellensburg, Washington is home to chimpanzees who use the signs of American Sign Language (ASL) to communicate with humans and each other. Four of the chimpanzees, Washoe, Moja, Tatu, and Dar were cross-fostered by humans and immersed in an ASL environment much like a child is immersed in a spoken language environment (Gardner & Gardner, 1989). The youngest chimpanzee, Loulis, was adopted and raised by Washoe. Loulis acquired his signs from Washoe and other signing chimpanzees (Fouts, Fouts, & Van Cantfort, 1989)....
... Bodamer, M.D. & Gardner, R.A. (2002). How cross-fostered chimpanzees (Pan troglodytes) initiate and maintain conversations. Journal of Comparative Psychology, 116, 12-26.
Fouts, R.S., Fouts, D.H., & Van Cantfort, T.E. (1989). The infant Loulis learns signs from cross-fostered chimpanzees. In R.A. Gardner, B.T. Gardner, & T.E. Van Cantfort (Eds.), Teaching Sign Language to Chimpanzees (pp. 280-292).
Jensvold, M. L. A., & Gardner, R. A. (2000). Interactive use of sign language by cross-fostered chimpanzees (Pan troglodytes). Journal of Comparative Psychology, 114, 335-346.