The following is a quote of Gould that previously was posted by Paul.
I added the bold face to two of the sentences. In the first bold face sentence, Gould is speaking about mutations. It is clear from the mathematics from ev that the larger the population, the more rapid the evolution (at least for random point mutations). However, if you consider a small population with advantageous alleles and recombination rather than mutations, Gould’s postulate makes sense.
The second bold faced sentence also makes sense if you are talking about recombination and natural selection. Pygmy elephants could rapidly evolve to their larger relatives that we see today by recombination and natural selection without their being much evidence in the fossil record because of the rapid transition from the small form to the large form. If you consider the analogy of dog breeding, you have the variation in size from Chihuahuas to Great Danes in fewer than 10,000 generations.
Gould should have been using the word recombination, not mutation in his postulate and then his theory would match observations.
Are Chihuahuas and Great Danes still members of the same species? I can't imagine that it would be reasonably likely for the two "breeds" to be able procreate without human intervention.